In the year 1962, the Radcliffe family and their one hundred and one dalmatians are preparing to move to their “Dalmatian Plantation,” a home in the countryside with plenty of room for all of them. However, one of the puppies, Patch, feels ignored and wishes to be unique like his television hero, Thunderbolt. While watching the Thunderbolt Adventure Hour, he hears about a chance to appear on the show. He is accidentally left behind when his family leaves for the plantation, so he decides to head for the audition to meet his hero and win a guest spot on the show. He fails to impress the producers, allowing a jealous Lil’ Lightning to manipulate them into making himself the hero of his own show. Depressed, Patch leaves and runs into Thunderbolt. Elsewhere in London, Cruella de Vil has returned, but she is under probation and a restraining order for her past behavior. No longer allowed to maintain her previous lifestyle, she attempts to soothe her fixation on spots with the help of Lars, a German artist. In order to inspire him, she restarts her hunt for the dalmatians, using a newspaper picture of Patch to find their new address. Patch’s family finally becomes aware that he is missing and go back to London to find him. Cruella bails her former henchmen, Jasper and Horace, out of prison. She sends them in a stolen dog food truck to steal the remaining puppies. They succeed after dealing with Nanny, and they take them to Lars. When Cruella requests she be made a masterpiece from their fur, Lars refuses, not wanting them to be harmed. Angered, she has him bound and gagged and returns to her original plan of making a dalmatian fur coat.